Geos 308 Discussion Assignment 4

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University Of Arizona *

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308

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Anthropology

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Dec 6, 2023

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For this assignment, you will read the following paper (found in the discussion readings folder): Carr, T. D., Napoli, J. G., Brusatte, S. L., Holtz, T. R., Hone, D. W., Williamson, T. E., & Zanno, L. E. (2022). Insufficient Evidence for Multiple Species of Tyrannosaurus in the Latest Cretaceous of North America: A Comment on “The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus Tyrannosaurus ”. Evolutionary Biology , 49 (3), 327-341. For some context on this paper, check out the NY Times coverage here: https://www.nytimes.com/2022/02/28/science/tyrannosaurus-rex-species.html https://www.nytimes.com/2022/07/25/science/tyrannosaurus-rex-dinosaurs.html Based on this paper, answer the following questions and bring them to class with you. 1. How did Paul et al. split Tyrannosaurus rex into three species? - Paul et al. claimed the species to be stratigraphically separate. 2. Paul et al. define two chronospecies and one morphospecies. What are these and what lines of evidence do they give to justify these species? - Two chronospecies being T. imperator > T. rex ) and one morphological species ( T. regina ) - There were three main lines of evidence given: 1) The skeletal robusticity was consistent with the specimen in question. 2) Tyrannosaurus femoral proportions do show unusual variation, (Paul et al, 2022). 3)the incisor teeth arrangement had a close correlation with femoral robusticity, (Paul et al, 2022). 3. How did Paul et al. diagnose the three species? Do you think these are sufficient diagnoses? - There was a procedure formed to be able to test the hypothesis that a fossil species may be several. First there needs to be a null and alternative hypotheses, from here a there needs to complete a quantitative cladistic analysis in order for it to be tested and observed if there are more than one species present. It is also mentioned for a agglomerative hierarchical test to be done so that the statistical analysis can show any differences in the data. The following approach from here is to obtain precise stratigraphic data which would distinguish chronospeices form morphological species. 4. What was the basis of Carr et al.’s rebuttal?
- The rebuttal lies in the basis that variation in the T.rex hypodigm is expectional, meaning that there are “robust” and “gracile” morph cryptic species present that do not belong solely to T.rex bur rather, to other species. 5. How did Carr et al. use the bird data to support their point? - A ‘density vs. range of instraspecific variation in femur robusticity’ histogram was made to how the variation correspondence of the T.rex and that of living birds today. 6. What is the problem, according to Carr et al., of using privately held specimens? - The problem with privately held specimens is an ethical standards issue due to
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