When did gills first appear?

Aquaculture Science
3rd Edition
ISBN:9781133558347
Author:Parker
Publisher:Parker
Chapter2: Aquatic Plants And Animals
Section: Chapter Questions
Problem 9E
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When did gills first appear?

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Introduction:

Gill is found in several aquatic living organisms and acts as their respiratory organ. It draws in the dissolved oxygen present in the water and gives off the carbon dioxide. 

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