What 4 populations were being studied and compared?

Biology: The Dynamic Science (MindTap Course List)
4th Edition
ISBN:9781305389892
Author:Peter J. Russell, Paul E. Hertz, Beverly McMillan
Publisher:Peter J. Russell, Paul E. Hertz, Beverly McMillan
Chapter56: Animal Behavior
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In the article “ Environmental and Social Determinants of Anuran Lekking Behavior: Intraspecific Variation in Populations at Thermal Extremes" 

What 4 populations were being studied and compared?

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