We might expect that populations near the periphery of a species’ range (marginal populations) might evolve greater tolerance of local conditions, which allow them to increase in numbers and thereby expand the species’ range. Why might this not be the case?

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We might expect that populations near the periphery of a species’ range (marginal
populations) might evolve greater tolerance of local conditions, which allow them to increase in
numbers and thereby expand the species’ range. Why might this not be the case?

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